Award-winning photography of natural history, landscape, antiquity and travel throughout the world. All images © Martin Grace
CATEGORY WINNER IN BIRD PHOTOGRAPHER OF THE YEAR

Portfolio - Wings

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A Mallard (Anas platyrhynchos) bathing, photographed contre jour, Leighton Moss (RSPB), England

Sub-adult (4th-year) Northern Gannet (Morus bassanus) in flight, about to land, and being watched by a Western Jackdaw (Corvus monedula), Bempton Cliffs (RSPB), East Riding of Yorkshire, England

Grey Heron (Ardea cinerea) in flight, Leighton Moss (RSPB), Carnforth, Lancashire, England

A Mute Swan (Cygnus olor) preening in angled, late afternoon sunlight

Female Grey Phalarope (Red Phalarope) (Phalaropus fulicarius) taking off from the surface of water, with floating kelp

Flying Nazca Booby (Sula granti) about to land, Punta Suarez, Espanola, Galapagos

Grey Heron (Ardea cinerea) in flight, Leighton Moss (RSPB), Carnforth, Lancashire, England

Black-Headed Gull (Larus ridibundus) in flight, Leighton Moss, Lancashire, England

Male Eider (Somateria mollissima) in flight, Holmiabukta, Svalbard, Norway

Swallow-Tailed Gull (Larus furcatus), Islote Pitt, San Cristobal, Galapagos Islands

Great Blue Heron (Ardea herodias) in flight, about to land, Bahia Ballena, Santa Cruz, Galapagos

Pied Kingfisher (Ceryle rudis), River Chobe, Botswana

Global Warming - Arctic Tern (Sterna paradisaea) at Longyearbyen, Svalbard, Norway

Pied Kingfisher (Ceryle rudis), River Chobe, Botswana

Great Frigatebird male (Fregata minor ridgwayi) flying with inflated gular pouch to attract a mate, Darwin Beach, Genovesa, Galapagos Islands

Immature African Jacana (Actophilornis africana) taking off, River Chobe, Botswana

Little Egret (Egretta garzetta) in flight just after sunrise, River Chobe, Botswana

Martial Eagle (Polemaetus bellicosus) just about to take off carrying prey Helmeted Guineafowl (Numida meleagris), Etosha, Namibia

Magnificent Frigatebird female (Fregata magnificens) harries a juvenile male Great Frigatebird (Fregata minor ridgwayi), Cerro Brujo, San Cristobal, Galapagos Islands

Clumsy landing - Black-Headed Gull (Larus ridibundus)

Image 1 of an extraordinary sequence of 6 images of a Hood Mockingbird (Nesomimus macdonaldi) running up the steep support wire for a beacon, Punta Suarez, Espanola, Galapagos

Juvenile Striated Heron (Butorides striatus) flying to escape a wave, Bahia Ballena, Santa Cruz, Galapagos

Black-Headed Gull (Larus ridibundus) in flight, Leighton Moss, Lancashire, England

Aggressive display from a Nazca Booby (Sula granti) towards another flying a little too low overhead, Punta Suarez, Espanola, Galapagos

Grey Heron (Ardea cinerea) in flight, Leighton Moss (RSPB), Carnforth, Lancashire, England

Little Egret (Egretta garzetta garzetta) taking off, Leighton Moss, Carnforth, Lancashire, England

African Jacana (Actophilornis africana) in the early morning sun, River Chobe, Botswana

Martial Eagle (Polemaetus bellicosus) in flight, carrying prey Helmeted Guineafowl (Numida meleagris), Etosha, Namibia

Silhouetted flying Pink-Footed Goose (Anas brachyrhynchos) preparing to land, Lancashire, England

Black-Headed Gull (Larus ridibundus) in flight

European Reed Warbler (Acrocephalus scirpaceus) in flight through reed bed, Parque Nacional de las Tablas de Daimiel, Castilla-La Mancha

Five Little Egrets (Egretta garzetta garzetta) - one taking off, Leighton Moss, Carnforth, Lancashire, England

Sub-adult (4th-year) Northern Gannet (Morus bassanus) in flight over others on a cliff edge, Bempton Cliffs (RSPB), East Riding of Yorkshire, England

Pied Kingfisher (Ceryle rudis), River Chobe, Botswana

Male African Darter or Anhinga (Anhinga rufa (melanogaster)) in breeding plumage at sunrise, River Chobe, Botswana

Common Shelduck (Tadorna tadorna) flying into late afternoon sunlight, Lancashire, England

Grey Heron (Ardea cinerea) in flight, Leighton Moss (RSPB), Carnforth, Lancashire, England

Little Egret (Egretta garzetta) in flight, River Chobe, Botswana

Black-Headed Gull (Larus ridibundus) in flight, Leighton Moss, Lancashire, England

Fast flight of four Black-Tailed Godwit (Limosa limosa) with background blurred, Dumfries & Galloway, Scotland

Common Gull (Mew Gull) (Larus canus) about to land on a paling fence, Lochindorb, Scottish Highlands

Antarctic Shag (Phalacrocorax (atriceps) bransfieldensis) in flight, Paulet Island, Antarctica

Antarctic Shag (Phalacrocorax (atriceps) bransfieldensis), Paulet Island, Antarctica

She may only be a Mallard (Anas platyrhynchos) but this balletic beauty is worthy of Swan Lake! Leighton Moss (RSPB), England

Gentoo Penguin (Pygoscelis papua) bathing in meltwater slush pool, Petermann Island, Antarctic Peninsula

Little Egret (Egretta garzetta) flying at sunset, River Chobe, Botswana

Black-Winged Stilt (Himantopus himantopus) in flight, Fuente de Piedra, Andalucia, Spain

Grey Heron (Ardea cinerea) in flight, Leighton Moss (RSPB), Carnforth, Lancashire, England

Cropped version of the previous image of an Alpine or Yellow-Billed Chough (Pyrrhocorax graculus) in flight, Picos de Europa, Cantabria, Spain

Bee-Fly (Bombylius major) hovering while it feeds from Cuckoo-flower (Cardamine pratensis)