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The dominant female Giant River Otter (Pteronura brasiliensis) defaecates into the communal latrine, River Piquiri, Northern Pantanal, Mato Grosso, Brazil
She will then puddle the ground with her feet, and roll in the mixture of sand, urine and faeces: all part of scent-marking territorial behaviour
Photo ID: PAN_06651_Giant_River_Otter_(Pteronura_brasiliensis)_Piquirí_River_Northern_Pantanal_August_10_CrssEA
Photo size: 17.1 Mpixels (48.8 MB uncompressed) - 5099x3345 pixels (17x11.2 in / 43.2x28.3 cm at 300 ppi)
Photo keywords: animal, ariranha, bank, behaviour, biology, Brasil, brasilensis, Brazil, Carnivora, carnivore, communal, Cuiabá, defaecate, defecate, Giant River Otter, head, latrine, mammal, mark, Martin Grace, Mato Grosso, Matogrossense, mustelid, Mustelidae, native, natural history, nature, otter, Pantanal, Pantanal Matogrossense, Piquiri, Pteronura, rare, river, riverbank, scent-marking, slide, territorial, threatened, urinate, urination, urine, vegetation, water, wild, wilderness, zoology
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